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In a dithecous anther, each pollen sac contains 1000 MMC. What is the total number of pollen grain produced by the anther?

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Hint: A dithecous anther contains two theca or two lobes and each theca contains a pollen sac. So, it is also called tetrasporangiate. Microspore mother cells are diploid and each microspore mother cell divides to form four microspores. Out of four microspores, three of them degenerate and one forms pollen grain.

Complete answer:
The total number of pollen grains produced by an anther is 4000 as each pollen sac in a dithecous anther contains 1000 MMC. A dithecous or tetrasporangiate anther has four chambers or pollen sac. These pollen sacs contain microspore mother cells which give rise to four microspores. Out of these four haploid microspores, three degenerate and one becomes the pollen grain. So, as each pollen sac contains 1000 MMC, the total number of pollen grains produced by the anther would be 4000.
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- An anther consists of four layers- outer epidermis, endothecium, middle layers, and tapetum. Tapetum is responsible for the nourishment of pollen grains.
- Epidermis is the protective layer. Endothecium consists of an outer thin layer and an inner layer of alpha cellulosic fibers which is hygroscopic in nature. The middle layer is ephemeral and helps in the dehiscence of the anther.
- Tapetum contains enzymes and hormones. It is of two types- glandular and invasive. Glandular tapetum secretes pro ubisch bodies which in turn produces ubisch which produces sporopollenin.
Note: An anther gives rise to pollen grain which contains the male gamete. The pollen grain is covered by an outer layer of exine and an inner layer of intine. The exine is made up of sporopollenin which is resistant to every microbial action and remains as such in solid or sediments. Sporopollenin keeps the pollen non degraded for a long time. Pollen grain has germ pores where sporopollenin is not present.